Pollination, Pollinizers, and Pollinators in Horticultural Crops
- Introduction
- Pollination is one of the most important processes in the reproductive cycle of flowering plants.
- It directly influences fruit set, fruit quality, seed development, and crop yield in both annual and perennial horticultural crops.
- In many fruit crops, cross-pollination is essential for successful fertilization and fruit development.
- Hence, knowledge of pollination biology, pollinizers, and pollinators is crucial for scientific orchard management and productivity.
- Definition of Pollination
- Pollination is the transfer of pollen grains from the anther (male part) of a flower to the stigma (female part) of the same or another flower.
- It is the first essential step leading to fertilization and subsequent fruit and seed formation.
- Types of Pollination
Pollination is classified mainly into two types, based on the source of pollen.
- Self-Pollination (Autogamy) Transfer of pollen grains from the anther to the stigma of the same flower or another flower on the same plant.
Types:
- Autogamy – within the same flower (e.g., Tomato, Pea).
- Geitonogamy – between flowers on the same plant (e.g., Mango, Banana).
Characteristics:
- Genetic uniformity.
- No dependence on external agents.
- Less variability, but may cause inbreeding depression.
Examples: Peach, Apricot, Tomato, Brinjal, Chilli.
- Cross-Pollination (Allogamy); Transfer of pollen grains from the anther of one flower to the stigma of a different plant of the same species.
Characteristics:
- Promotes genetic variation.
- Ensures hybrid vigor (heterosis).
- Requires external agents (wind, insects, etc.).
- Common in most fruit trees.
Examples: Apple, Pear, Plum, Almond, Mango, Citrus.
- Types of Cross-Pollination Agents
Cross-pollination requires external agents for pollen transfer known as pollinating agents.
|
Mode of Pollination |
Agent |
Examples |
|
Anemophily |
Wind |
Coconut, Date palm, Walnuts |
|
Entomophily |
Insects (bees, butterflies, beetles) |
Apple, Citrus, Mango, Guava |
|
Hydrophily |
Water |
Rare in horticultural crops |
|
Zoophily |
Birds, bats, animals |
Banana (bats), Hummingbirds (in some ornamentals) |
- Importance of Pollination in Fruit Crops
|
Importance |
Explanation |
|
1. Fruit and seed set |
Pollination is essential for fertilization and seed development. |
|
2. Fruit quality |
Proper pollination ensures uniform fruit shape, size, and seed number. |
|
3. Yield |
Increases fruit set percentage and overall productivity. |
|
4. Hybridization |
Used in plant breeding for developing new varieties. |
|
5. Genetic diversity |
Maintains variability and adaptability in crops. |
Pollinizers
- Definition
- A pollinizer is a plant or cultivar that provides viable and compatible pollen for the fertilization of another plant’s flowers.
- Pollinizers are essential in self-incompatible or cross-pollinated crops.
- Characteristics of a Good Pollinizer
|
Characteristic |
Description |
|
1. Genetic compatibility |
Must be of the same species but a different, compatible variety. |
|
2. Simultaneous flowering |
Flowering period must overlap with the main variety. |
|
3. High pollen viability and fertility |
Pollen must germinate effectively on stigma. |
|
4. Abundant pollen production |
Produces large amounts of viable pollen. |
|
5. Similar climatic adaptability |
Should thrive under the same growing conditions. |
Importance of Pollinizers
- Ensures effective cross-pollination and fertilization.
- Improves fruit set percentage and quality.
- Maintains regular bearing and prevents unfruitfulness.
- Used for hybrid seed production and breeding programs.
Examples of Pollinizers in Major Fruit Crops
|
Main Variety |
Pollinizer Variety |
Remarks |
|
Apple (Delicious group) |
Golden Delicious, Granny Smith |
Self-incompatible, needs cross-pollination. |
|
Pear |
Bartlett, Kieffer |
Requires compatible pollinizer. |
|
Plum |
Santa Rosa, Wickson |
Cross-compatible varieties required. |
|
Almond |
Nonpareil × Peerless |
Cross-pollinated by bees. |
|
Mango (Dashehari) |
Langra, Chausa, Amrapali |
Mixed planting improves fruit set. |
|
Citrus (Sweet orange) |
Mandarins or other Citrus species |
Cross-compatibility increases fruit set. |
|
Litchi |
Bombai, Bedana |
Mixed varieties increase pollination. |
Pollinizer Planting Design in Orchards
- Pollinizers should be planted systematically to ensure adequate pollen transfer.
|
Method |
Description |
|
Block planting |
Separate rows or blocks of pollinizers (used in apple, pear). |
|
Alternate planting |
Every 3rd or 4th tree is a pollinizer. |
|
Interplanting |
Pollinizer trees mixed within the main variety. |
|
Ratio |
Generally 1 pollinizer : 9–10 main trees (10%). |
Pollinators
- Definition
- Pollinators are living agents (mostly insects, birds, or animals) that carry pollen from the anther of one flower to the stigma of another, resulting in pollination.
- They play a vital role in cross-pollinated and insect-pollinated crops.
- Types of Pollinators
|
Pollinator Type |
Examples |
Pollination Type |
|
Insects |
Honeybees, Butterflies, Beetles |
Entomophily |
|
Birds |
Sunbirds, Hummingbirds |
Ornithophily |
|
Bats |
Fruit bats |
Chiropterophily |
|
Wind |
None (mechanical agent) |
Anemophily |
|
Humans |
Artificial hand pollination |
Manual pollination |
- Importance of Pollinators
|
Benefit |
Explanation |
|
1. Enhanced fruit set |
Essential for self-incompatible crops (apple, pear, plum). |
|
2. Better fruit quality |
Ensures uniform fertilization and seed distribution. |
|
3. Increased yield |
More pollination → higher productivity. |
|
4. Regular bearing |
Reduces unfruitfulness and irregular flowering. |
|
5. Pollination in hybrid breeding |
Essential for controlled hybridization. |
- Honeybees as Major Pollinators
- The most efficient natural pollinators in horticultural crops.
- Bees visit flowers for nectar and pollen and transfer pollen between flowers during their visits.
Advantages:
- Promote cross-pollination and fertilization.
- Improve fruit shape, size, and seed content.
- Increase yield up to 30–50% in crops like apple, citrus, and cucurbits.
Management Practices:
- Introduce 2–3 honeybee colonies per hectare during flowering.
- Avoid pesticide spraying during bloom.
- Ensure flowering overlap between varieties.
- Artificial Pollination (Hand Pollination)
When natural pollinators are absent or insufficient, manual pollination is practiced.
|
Crop |
Method |
Purpose |
|
Date palm |
Hand pollination using male inflorescence |
Ensures fruit set |
|
Custard apple |
Manual pollen transfer using brush |
Improves fruit set |
|
Vanilla |
Artificial pollination with stick |
Essential for fruiting |
|
Apple / Pear (Controlled breeding) |
Controlled cross-pollination |
Hybridization |
- Pollination Management in Orchards
|
Practice |
Objective |
|
Plant compatible pollinizers |
Ensure cross-pollination. |
|
Maintain bee colonies |
Promote natural pollination. |
|
Avoid pesticides during bloom |
Protect pollinators. |
|
Provide flowering diversity |
Attract and sustain pollinators. |
|
Maintain windbreaks and shelterbelts |
Protect bees and flowers from wind. |
- Factors Affecting Pollination Success
|
Factor |
Effect |
|
Weather |
Rain, wind, or cold reduce bee activity and pollen viability. |
|
Time of day |
Morning hours are ideal for pollination (maximum bee activity). |
|
Pollen viability |
Declines with age or under high humidity. |
|
Flower morphology |
Inaccessible or closed flowers hinder pollination. |
|
Pesticide use |
Harmful to bees and other pollinators. |
|
Flowering synchronization |
Must overlap between pollinizer and main variety. |
- Examples of Pollination Systems in Fruit Crops
|
Crop |
Type of Pollination |
Pollinator / Agent |
|
Apple |
Cross-pollination |
Bees, wind |
|
Mango |
Cross-pollination |
Insects (flies, bees) |
|
Citrus |
Cross/self-pollination |
Bees |
|
Guava |
Mainly self, sometimes cross |
Insects |
|
Papaya |
Cross-pollination (dioecious) |
Wind, insects |
|
Banana |
Parthenocarpic (no pollination needed) |
None |
|
Date palm |
Dioecious, artificial pollination |
Hand pollination |
|
Grapes |
Self-pollination |
Wind, insects |
|
Pomegranate |
Cross/self |
Bees |
